Document Type : Research Paper

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Abstract

One-year seedlings of five species of eucalypt: E. camaldulensis, E. microtheca, E. viminalis, E. globulus and E. gunnii were treated for 14 weeks in growth season, tilting a branch to c. 60° in every successful ones. We investigated the anatomical features of stems wood after 15 days and every 2 weeks according to IAWA list of features afterwards.
The results showed that tilted stems formed more tension wood in the form of gelatinous fibers than straight (untreated) ones. The formation of gelatinous fibers was considered to be closely related to tension wood formation. The formation of gelatinous fibers was considered to be closely related to tension wood formation. Excluding E. globulus, all the other species formed growth eccentricity and tension wood. Although the significant differences between the species and between different locations in the branches, there were no significant differences between the percent of tension wood among the species studied.

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-Aguayo, M.G., Mendonça, R.T., Martínez, P., Rodríguez, R. and Pereira, M., 2012. Chemical characteristics and Kraft pulping of tension wood from Eucalyptus globulus labill, Revista Árvore,36(6): 1163-1172

-Badia, M.A., Constant, T., Mothe, F. and Nepveu, G., 2006. Tension wood occurrence in three cultivars of Populus × euramericana. Part I: Inter-clonal and intra-tree variability of tension wood, Ann. For. Sci. 63: 23–30
-Bailleres, H., Fournier,  B., Tollier, M., Monties, M. T. and Chanson B., 1995. Structure, composition chimique et retraits de maturation du bois chez les clones d’Eucalyptus. Ann. Sci. For. 52: 157-172
-Bamber, R. K., 2001. A general theory for the origin of growth stresses in reaction wood: How trees stay upright. IAWA Journal, 22 (3): 205-212.
-Berlyn, G. P., 1961. Factors affecting the incidence of reaction tissue in Populus deltoides,Iowa state J. Sci. 35 (3): 367-424.
-Chafe, S. C., 1977. Radial dislocations in the fiber wall of Eucalyptus regnans trees of high growth stress. Wood Sci. Technol. 11: 69-77.
-Chudnoff, M. and Tischler K., 1963. Fiber morphology of Eucalyptus camaldulensis Dehn. Suppl. No. 1, PP. 23.
-Fahn, A., 1990. Plant anatomy. 4th ed. Pergamon press, Oxford, PP. 588
-Hoseinzadeh, j., Abdolkhani, A., Emaminasab, M., Khodabandelou, H. and Ahmadi, M., 2015. Characteristics of paper made from poplar tension wood and normal wood, Iranian Journal Wood Paper Research, 30(4):76-78
-IAWA Committee, 1989. IAWA list of microscopic features for hardwood identification. IAWA Bul. n.s. 10(3):219-261
-Isebrands, J. G. & Bensend, D.W., 1972. Incidence and structure of gelatinous fibers within rapid-growing eastern cottonwood. Wood Fiber 4 (2) 61-71.
-Jourez, B., Riboux, A. and Leclercq A., 2001. Anatomical characteristics of tension Wood and opposite Wood in young inclined stems of poplar (Populus euramericana cv."Ghoy"), IAWA Journal, Vol. 22 (2): 133-157.
-Onaka, F., 1949. Studies on compression and tension Wood. Wood Research Bulletin, 1:1–88.
-Ruelle, J., Yoshida, M., Clair, B. and Thibaut, B.,  2007. Peculiar tension wood structure in Laetia procera (Poepp.) Eichl.(Flacourtiaceae).Trees Structure and Function. 21 (3): 345-355.
- Sayar, M. and Tarmian A., 2010.The effect of tension wood formation on the anatomical characteristics and gelatinous layer formation in the fibers of poplar (P. nigra), J. of Wood & Forest Science and Technology, 17(2):133-144
-Tarmian, A., Faezipour, M., Karimi, A., Parsapajouh, D. and Moezipour, A., 2008.Beech:comparing physical properties and fibre biometry between tension wood and normal wood. Pajouhesh va sazandegi ,21(4):39-45.
-Toghraie, N., 2009. Final  report of Formation and lignifications of xylem  tissue of tension wood and  normal  wood in 3 Eucalypt species, RIFR Pub. 113 p.
-Toghraie, N., Hedjazi, R., Parsapajouh, D. and Yazdani moghadam, H., 2007.The early formation of tension  wood  in Eucalyptus gunnii branches, Pajouhesh va Sazandegi, 76: 32-38
-Toghraie, N., Parsapajouh, D., Ebrahimzadeh, H., Thibaut, B., Gril, J., and  Yazdani moghadam, H., 2006. Tension wood in eucalypt trees. Journal of Science (University of Tehran), 32, 13-22

-Yoshida,  M., Okuda T. and Okuyama, T., 2000. Tension Wood and growth stress induced by artificial inclination in Liriodendron tulipifera Linn and Prunus sp. Annals of Forest Science.. 57: 739-746.